Nathan Lo, MD, PhD

Faculty Fellow in Infectious Diseases
Division of HIV, Infectious Diseases, and Global Medicine
Department of Medicine

Nathan C. Lo, MD PhD is a Faculty Fellow in Infectious Diseases (Research) in the Division of  HIV, Infectious Diseases, and Global Medicine at UCSF/ZSFG. He is a core faculty member in  the Experimental and Population-based Pathogen Investigation center (EPPIcenter), and a  clinical fellow in the Division of Infectious Diseases. He is an infectious diseases epidemiologist and public health scientist. The overarching goal of his research is to apply diverse computational methodologies to understand transmission of infectious diseases and design more effective public health policies to control and eliminate them locally and globally, with particular attention to vulnerable populations. His current research focuses on tropical diseases, vaccine-preventable infections, and COVID-19. He received a BS in Bioengineering from Rice University, and MD and PhD in Epidemiology from Stanford University. He completed a clinical residency in internal medicine at UCSF. He has been based at UCSF since 2019.


Review of 2022 WHO guidelines on the control and elimination of schistosomiasis.

The Lancet. Infectious diseases

Lo NC, Bezerra FSM, Colley DG, Fleming FM, Homeida M, Kabatereine N, Kabole FM, King CH, Mafe MA, Midzi N, Mutapi F, Mwanga JR, Ramzy RMR, Satrija F, Stothard JR, Traoré MS, Webster JP, Utzinger J, Zhou XN, Danso-Appiah A, Eusebi P, Loker ES, Obonyo CO, Quansah R, Liang S, Vaillant M, Murad MH, Hagan P, Garba A

Contribution and quality of mathematical modeling evidence in World Health Organization guidelines: A systematic review.


Lo NC, Andrejko K, Shukla P, Baker T, Sawin VI, Norris SL, Lewnard JA

COVID-19 Vaccination and Estimated Public Health Impact in California.

JAMA network open

Tan ST, Park HJ, Rodríguez-Barraquer I, Rutherford GW, Bibbins-Domingo K, Schechter R, Lo NC

Risk factor targeting for vaccine prioritization during the COVID-19 pandemic.

Scientific reports

Chapman LAC, Shukla P, Rodríguez-Barraquer I, Shete PB, León TM, Bibbins-Domingo K, Rutherford GW, Schechter R, Lo NC

Predicting covid-19 outcomes.

BMJ (Clinical research ed.)

Habib AR, Lo NC

Comparison of Strategies for Typhoid Conjugate Vaccine Introduction in India: A Cost-Effectiveness Modeling Study.

The Journal of infectious diseases

Ryckman T, Karthikeyan AS, Kumar D, Cao Y, Kang G, Goldhaber-Fiebert JD, John J, Lo NC, Andrews JR

Geographic Pattern of Typhoid Fever in India: A Model-Based Estimate of Cohort and Surveillance Data.

The Journal of infectious diseases

Cao Y, Karthikeyan AS, Ramanujam K, Raju R, Krishna S, Kumar D, Ryckman T, Mohan VR, Kang G, John J, Andrews JR, Lo NC

Frequency of Routine Testing for Coronavirus Disease 2019 (COVID-19) in High-risk Healthcare Environments to Reduce Outbreaks.

Clinical infectious diseases : an official publication of the Infectious Diseases Society of America

Addendum needed on COVID-19 travel study - Authors' reply.

The Lancet. Infectious diseases

Kiang MV, Chin ET, Huynh BQ, Chapman LAC, Lo NC

Comparison of infection control strategies to reduce COVID-19 outbreaks in homeless shelters in the United States: a simulation study.

BMC medicine

Chapman LAC, Kushel M, Cox SN, Scarborough A, Cawley C, Nguyen TQ, Rodriguez-Barraquer I, Greenhouse B, Imbert E, Lo NC

Routine asymptomatic testing strategies for airline travel during the COVID-19 pandemic: a simulation study.

The Lancet. Infectious diseases

Kiang MV, Chin ET, Huynh BQ, Chapman LAC, Rodríguez-Barraquer I, Greenhouse B, Rutherford GW, Bibbins-Domingo K, Havlir D, Basu S, Lo NC

Influenza, Varicella, and Mumps Outbreaks in US Migrant Detention Centers.


Lo NC, Nyathi S, Chapman LAC, Rodriguez-Barraquer I, Kushel M, Bibbins-Domingo K, Lewnard JA

Comparison of World Health Organization and Demographic and Health Surveys data to estimate sub-national deworming coverage in pre-school aged children.

PLoS neglected tropical diseases

Lo NC, Gupta R, Addiss DG, Bendavid E, Heft-Neal S, Mikhailov A, Montresor A, Mbabazi PS

Childhood vaccines and antibiotic use in low- and middle-income countries.


Lewnard JA, Lo NC, Arinaminpathy N, Frost I, Laxminarayan R

Hunter's Tropical Medicine and Emerging Infectious Diseases

Neglected Tropical Diseases: Public Health Control Programs and Mass Drug Administration

PJ Hotez, NC Lo

The 2016 California policy to eliminate nonmedical vaccine exemptions and changes in vaccine coverage: An empirical policy analysis.

PLoS medicine

Nyathi S, Karpel HC, Sainani KL, Maldonado Y, Hotez PJ, Bendavid E, Lo NC

Clinical evaluation for morbidity associated with soil-transmitted helminth infection in school-age children on Pemba Island, Tanzania.

PLoS neglected tropical diseases

Bogoch II, Speich B, Lo NC, Moser W, Croll D, Ali SM, Ame SM, Utzinger J, Andrews JR, Keiser J

Paediatric tuberculosis transmission outside the household: challenging historical paradigms to inform future public health strategies.

The Lancet. Respiratory medicine

Martinez L, Lo NC, Cords O, Hill PC, Khan P, Hatherill M, Mandalakas A, Kay A, Croda J, Horsburgh CR, Zar HJ, Andrews JR

Comparison of Strategies and Incidence Thresholds for Vi Conjugate Vaccines Against Typhoid Fever: A Cost-effectiveness Modeling Study.

The Journal of infectious diseases

Lo NC, Gupta R, Stanaway JD, Garrett DO, Bogoch II, Luby SP, Andrews JR

Modulation of Antibody Responses to the V1V2 and V3 Regions of HIV-1 Envelope by Immune Complex Vaccines.

Frontiers in immunology

Hioe CE, Kumar R, Upadhyay C, Jan M, Fox A, Itri V, Peachman KK, Rao M, Liu L, Lo NC, Tuen M, Jiang X, Kong XP, Zolla-Pazner S

Typhoid conjugate vaccines: a new tool in the fight against antimicrobial resistance.

The Lancet. Infectious diseases

Andrews JR, Baker S, Marks F, Alsan M, Garrett D, Gellin BG, Saha SK, Qamar FN, Yousafzai MT, Bogoch II, Antillon M, Pitzer VE, Kim JH, John J, Gauld J, Mogasale V, Ryan ET, Luby SP, Lo NC

Deworming in pre-school age children: A global empirical analysis of health outcomes.

PLoS neglected tropical diseases

Lo NC, Snyder J, Addiss DG, Heft-Neal S, Andrews JR, Bendavid E

Review of the 2017 WHO Guideline: Preventive chemotherapy to control soil-transmitted helminth infections in at-risk population groups. An opportunity lost in translation.

PLoS neglected tropical diseases

Savioli L, Albonico M, Daumerie D, Lo NC, Stothard JR, Asaolu S, Tchuem Tchuenté LA, Anderson RM

Impact and cost-effectiveness of snail control to achieve disease control targets for schistosomiasis.

Proceedings of the National Academy of Sciences of the United States of America

Lo NC, Gurarie D, Yoon N, Coulibaly JT, Bendavid E, Andrews JR, King CH

Schistosoma haematobium Egg Excretion does not Increase after Exercise: Implications for Diagnostic Testing.

The American journal of tropical medicine and hygiene

Coulibaly JT, Andrews JR, Lo NC, N'Goran EK, Utzinger J, Keiser J, Bogoch II

Mobile phone and handheld microscopes for public health applications.

The Lancet. Public health

Bogoch II, Lundin J, Lo NC, Andrews JR

Mobile-phone and handheld microscopy for neglected tropical diseases.

PLoS neglected tropical diseases

Rajchgot J, Coulibaly JT, Keiser J, Utzinger J, Lo NC, Mondry MK, Andrews JR, Bogoch II

A call to strengthen the global strategy against schistosomiasis and soil-transmitted helminthiasis: the time is now.

The Lancet. Infectious diseases

Lo NC, Addiss DG, Hotez PJ, King CH, Stothard JR, Evans DS, Colley DG, Lin W, Coulibaly JT, Bustinduy AL, Raso G, Bendavid E, Bogoch II, Fenwick A, Savioli L, Molyneux D, Utzinger J, Andrews JR

Comparison of sensitivity and faecal egg counts of Mini-FLOTAC using fixed stool samples and Kato-Katz technique for the diagnosis of Schistosoma mansoni and soil-transmitted helminths.

Acta tropica

Coulibaly JT, Ouattara M, Becker SL, Lo NC, Keiser J, N'Goran EK, Ianniello D, Rinaldi L, Cringoli G, Utzinger J

Evaluation of a Urine Pooling Strategy for the Rapid and Cost-Efficient Prevalence Classification of Schistosomiasis.

PLoS neglected tropical diseases

Lo NC, Coulibaly JT, Bendavid E, N'Goran EK, Utzinger J, Keiser J, Bogoch II, Andrews JR

Improving helminth treatment access: costs and opportunities.

The Lancet. Infectious diseases

Lo NC, Andrews JR, Bogoch II

Assessment of global guidelines for preventive chemotherapy against schistosomiasis and soil-transmitted helminthiasis: a cost-effectiveness modelling study.

The Lancet. Infectious diseases

Lo NC, Lai YS, Karagiannis-Voules DA, Bogoch II, Coulibaly JT, Bendavid E, Utzinger J, Vounatsou P, Andrews JR

Cost-effectiveness of community-wide treatment for helminthiasis--Authors' reply.

The Lancet. Global health

Lo NC, Bogoch II, Utzinger J, Andrews JR

Comparison of community-wide, integrated mass drug administration strategies for schistosomiasis and soil-transmitted helminthiasis: a cost-effectiveness modelling study.

The Lancet. Global health

Lo NC, Bogoch II, Blackburn BG, Raso G, N'Goran EK, Coulibaly JT, Becker SL, Abrams HB, Utzinger J, Andrews JR

Strategies for last mile implementation of global health technologies.

The Lancet. Global health

Chao TE, Lo NC, Mody GN, Sinha SR

Interaction of Shiga toxin with the A-domains and multimers of von Willebrand Factor.

The Journal of biological chemistry

Lo NC, Turner NA, Cruz MA, Moake J

The aging of biomedical research in the United States.

PloS one

Matthews KR, Calhoun KM, Lo N, Ho V